FIG. 1. — Geological map showing the desert in southwestern Morocco and the location of the Sabkha of Gueran relative to Boujdour city. Red symbols mark sites that yielded vertebrates remains described herein. Inset shows the geographical distribution of Eocene marine faunal localities on the African continent. The main African localities of interest are numbered from 1 to 8. Numbers: 1, Gueran locality in southwestern Morocco, Bartonian (Gingerich & Zouhri 2015; Zouhri et al. 2018; 2, Garitas and nearby sites south of Ad-Dakhla in southwestern Morocco Bartonian-Priabonian (Zouhri et al. 2014, 2018; Gingerich & Zouhri 2017); 3, Ndomor Diop site near Taïba Ndiaye in Senegal, Lutetian (Hautier et al. 2014); 4, Tiavandou in Senegal, Priabonian (Elouard 1981); 5, Kpogame in Togo, Lutetian (Gingerich & Cappetta 2014); 6, Ameke in southern Nigeria, Bartonian (Andrews 1920; Halstead & Middleton 1974, 1976); 7, Fayum Depression in Egypt, which yielded a rich vertebrate faunas of Eocene-Oligocene age; 8, Dur At-Talah in Sirt Basin in Libya, Eocene-Oligocene (Abouessa et al. 2012). The geology is from Hollard et al. (1985) and map modified from Gingerich & Zouhri (2015).