FIGURE 6 Species-level variation in avian functional traits in relation to geography and lifestyle. Hand-wing index (wing elongation) peaks towards high latitudes (a), and in species with aquatic and aerial lifestyles (b); relative tarsus length peaks at mid-latitudes and non-forest regions (c), and in species with terrestrial lifestyles (d); relative beak length peaks in the tropics, including rainforests (e), and in nectar feeders and aquatic predators (f). For maps, median trait values were calculated for 18,709 grid-cell assemblages worldwide. Darker colours indicate larger trait values. Assemblages were delimited by extracting species native resident or breeding distributions (n = 10,964 species for which both trait and geographical range data are available) onto an equal area grid with a cell resolution of ~100 km (Behrmann projection). Relative beak and tarsus length are the residuals of a linear regression of log-transformed tarsus and beak length (mm) against log-transformed body mass (grams). Species in (b,d) are classified according to primary lifestyle (predominant locomotory niche; insessorial = perching lifestyle). Species in (f) are classified according to primary diet following Pigot et al. (2020). Sample sizes (b,d,f) are numbers of species in each category